If grazing pressure does not allow the development of scrub or
woodland on the fixed dunes, the fixed dune grasslands will eventually
develop into a different vegetation type. The surface of the soil
continues to be leached by rainwater and this gradually washes out
the basic minerals (especially calcium) which caused the high pH
of the earlier successional stages.
Without a large plant biomass to lock up the plant nutrients, these
are also leached out of the soil surface. The result is a dry, acid,
nutrient-poor substrate. Plants which are adapted to these conditions
are those of acid grasslands and heathlands, and on very old parts
of the dune system these vegetation types can develop. |